A New Taxon of Basal Ceratopsian from China and the Early Evolution of Ceratopsia
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{"title"=>"A new taxon of basal ceratopsian from China and the early evolution of Ceratopsia", "type"=>"journal", "authors"=>[{"first_name"=>"Fenglu", "last_name"=>"Han", "scopus_author_id"=>"25923913300"}, {"first_name"=>"Catherine A.", "last_name"=>"Forster", "scopus_author_id"=>"7201641486"}, {"first_name"=>"James M.", "last_name"=>"Clark", "scopus_author_id"=>"7407825960"}, {"first_name"=>"Xing", "last_name"=>"Xu", "scopus_author_id"=>"7405294877"}], "year"=>2015, "source"=>"PLoS ONE", "identifiers"=>{"isbn"=>"0028-0836 (Print)", "pmid"=>"26649770", "doi"=>"10.1371/journal.pone.0143369", "pui"=>"608000928", "issn"=>"19326203", "sgr"=>"84956669534", "scopus"=>"2-s2.0-84956669534"}, "id"=>"c1a1960d-2b3f-39f7-9ab4-656483a51c6b", "abstract"=>"Ceratopsia is one of the best studied herbivorous ornithischian clades, but the early evolution of Ceratopsia, including the placement of Psittacosaurus, is still controversial and unclear. Here, we report a second basal ceratopsian, Hualianceratops wucaiwanensis gen. et sp. nov., from the Upper Jurassic (Oxfordian) Shishugou Formation of the Junggar Basin, northwestern China. This new taxon is characterized by a prominent caudodorsal process on the subtemporal ramus of the jugal, a robust quadrate with an expansive quadratojugal facet, a prominent notch near the ventral region of the quadrate, a deep and short dentary, and strongly rugose texturing on the lateral surface of the dentary. Hualianceratops shares several derived characters with both Psittacosaurus and the basal ceratopsians Yinlong, Chaoyangsaurus, and Xuanhuaceratops. A new comprehensive phylogeny of ceratopsians weakly supports both Yinlong and Hualianceratops as chaoyangsaurids (along with Chaoyangsaurus and Xuanhuaceratops), as well as the monophyly of Chaoyangosauridae + Psittacosaurus. This analysis also weakly supports the novel hypothesis that Chaoyangsauridae + Psittacosaurus is the sister group to the rest of Neoceratopsia, suggesting a basal split between these clades before the Late Jurassic. This phylogeny and the earliest Late Jurassic age of Yinlong and Hualianceratops imply that at least five ceratopsian lineages (Yinlong, Hualianceratops, Chaoyangsaurus + Xuanhuaceratops, Psittacosaurus, Neoceratopsia) were present at the beginning of the Late Jurassic.", "link"=>"http://www.mendeley.com/research/new-taxon-basal-ceratopsian-china-early-evolution-ceratopsia", "reader_count"=>18, "reader_count_by_academic_status"=>{"Professor > Associate Professor"=>1, "Researcher"=>7, "Student > Doctoral Student"=>1, "Student > Ph. D. Student"=>1, "Student > Master"=>1, "Other"=>2, "Student > Bachelor"=>5}, "reader_count_by_user_role"=>{"Professor > Associate Professor"=>1, "Researcher"=>7, "Student > Doctoral Student"=>1, "Student > Ph. D. Student"=>1, "Student > Master"=>1, "Other"=>2, "Student > Bachelor"=>5}, "reader_count_by_subject_area"=>{"Unspecified"=>1, "Biochemistry, Genetics and Molecular Biology"=>1, "Agricultural and Biological Sciences"=>6, "Earth and Planetary Sciences"=>10}, "reader_count_by_subdiscipline"=>{"Earth and Planetary Sciences"=>{"Earth and Planetary Sciences"=>10}, "Agricultural and Biological Sciences"=>{"Agricultural and Biological Sciences"=>6}, "Biochemistry, Genetics and Molecular Biology"=>{"Biochemistry, Genetics and Molecular Biology"=>1}, "Unspecified"=>{"Unspecified"=>1}}, "reader_count_by_country"=>{"Canada"=>1, "Japan"=>1}, "group_count"=>1}

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  • {"files"=>["https://ndownloader.figshare.com/files/2606252", "https://ndownloader.figshare.com/files/2606253"], "description"=>"<div><p>Ceratopsia is one of the best studied herbivorous ornithischian clades, but the early evolution of Ceratopsia, including the placement of <i>Psittacosaurus</i>, is still controversial and unclear. Here, we report a second basal ceratopsian, <i>Hualianceratops wucaiwanensis</i> gen. et sp. nov., from the Upper Jurassic (Oxfordian) Shishugou Formation of the Junggar Basin, northwestern China. This new taxon is characterized by a prominent caudodorsal process on the subtemporal ramus of the jugal, a robust quadrate with an expansive quadratojugal facet, a prominent notch near the ventral region of the quadrate, a deep and short dentary, and strongly rugose texturing on the lateral surface of the dentary. <i>Hualianceratops</i> shares several derived characters with both <i>Psittacosaurus</i> and the basal ceratopsians <i>Yinlong</i>, <i>Chaoyangsaurus</i>, and <i>Xuanhuaceratops</i>. A new comprehensive phylogeny of ceratopsians weakly supports both <i>Yinlong</i> and <i>Hualianceratops</i> as chaoyangsaurids (along with <i>Chaoyangsaurus</i> and <i>Xuanhuaceratops</i>), as well as the monophyly of Chaoyangosauridae + <i>Psittacosaurus</i>. This analysis also weakly supports the novel hypothesis that Chaoyangsauridae + <i>Psittacosaurus</i> is the sister group to the rest of Neoceratopsia, suggesting a basal split between these clades before the Late Jurassic. This phylogeny and the earliest Late Jurassic age of <i>Yinlong</i> and <i>Hualianceratops</i> imply that at least five ceratopsian lineages (<i>Yinlong</i>, <i>Hualianceratops</i>, <i>Chaoyangsaurus</i> + <i>Xuanhuaceratops</i>, <i>Psittacosaurus</i>, Neoceratopsia) were present at the beginning of the Late Jurassic.</p></div>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623040, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>["https://dx.doi.org/10.1371/journal.pone.0143369.s001", "https://dx.doi.org/10.1371/journal.pone.0143369.s002"], "stats"=>{"downloads"=>1, "page_views"=>2, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_A_New_Taxon_of_Basal_Ceratopsian_from_China_and_the_Early_Evolution_of_Ceratopsia_/1623040", "title"=>"A New Taxon of Basal Ceratopsian from China and the Early Evolution of Ceratopsia", "pos_in_sequence"=>0, "defined_type"=>4, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606226"], "description"=>"<p>(A) photograph in lateral view, (B) photograph in medial view, (C) drawing in lateral view, (D) drawing in medial view. Abbreviations: afo, antorbital fossa; cp, caudomedial process of the jugal; dp, caudodorsal process of the jugal; j, jugal; jn, jugal node; ma, maxilla; or, orbit; po, postorbital; sq, squamosal.</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623018, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.g001", "stats"=>{"downloads"=>0, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_The_articulated_right_maxilla_and_jugal_of_Hualianceratops_wucaiwanensis_IVPP_V18641_/1623018", "title"=>"The articulated right maxilla and jugal of <i>Hualianceratops wucaiwanensis</i> (IVPP V18641).", "pos_in_sequence"=>0, "defined_type"=>1, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606232"], "description"=>"<p>(A) photograph in lateral view, (B) drawing in lateral view, (C) photograph in ventral view with inset of a tooth enlarged, (D) drawing in ventral view, (E) photograph in dorsal view, (F) drawing in dorsal view, (G) photograph in medial view, (H) drawing in medial view. Abbreviations: ma, maxilla; j, jugal.</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623024, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.g002", "stats"=>{"downloads"=>0, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_The_articulated_right_maxilla_and_partial_jugal_of_Hualianceratops_wucaiwanensis_IVPP_V18641_the_broken_rostral_part_of_the_elements_in_Fig_1_/1623024", "title"=>"The articulated right maxilla and partial jugal of <i>Hualianceratops wucaiwanensis</i> (IVPP V18641), the broken rostral part of the elements in Fig 1.", "pos_in_sequence"=>0, "defined_type"=>1, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606235"], "description"=>"<p>(A) photograph in lateral view, (B) photograph in medial view, (C) drawing in lateral view; (D) drawing in medial view. Abbreviations: cp, caudomedial process of the jugal; dp, caudodorsal process of the jugal; qj, quadratojugal; q, quadrate; j, jugal.</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623027, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.g003", "stats"=>{"downloads"=>0, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_The_articulated_left_partial_jugal_and_quadratojugal_of_Hualianceratops_wucaiwanensis_IVPP_V18641_/1623027", "title"=>"The articulated left partial jugal and quadratojugal of <i>Hualianceratops wucaiwanensis</i> (IVPP V18641).", "pos_in_sequence"=>0, "defined_type"=>1, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606243"], "description"=>"<p>Outline drawing of the holotype skull of <i>Yinlong downsi</i> showing the infratemporal ramus of the jugal (red) in lateral view (A) and ventral view (B), and the infratemporal ramus in basal ceratopsians in right lateral view (C-H) and ventral view (I-N). (C) <i>Hualianceratops wucaiwanensis</i> (IVPP V18641), (D) <i>Yinlong downsi</i> (IVPP V14530), (E) <i>Hongshanosaurus houi</i> (IVPP V12617), (F) <i>Chaoyangsaurus youngi</i> (IGCAGS V371), (G) <i>Liaoceratops yanzigouensis</i> (IVPP V12633), (H) <i>Archaeoceratops oshimai</i> (IVPP V11114). (I) <i>Hualianceratops wucaiwanensis</i> (IVPP V18641), (J) <i>Yinlong downsi</i> (IVPP V14530), (K) <i>Hongshanosaurus houi</i> (IVPP V12617), (L) <i>Chaoyangsaurus youngi</i> (IGCAGS V371), (M) <i>Liaoceratops yanzigouensis</i> (IVPP V12633), (N) <i>Archaeoceratops oshimai</i> (IVPP V11114). The arrows show the caudal process of the infratemporal ramus of the jugal. Abbreviations: j, jugal; qj, quadratojugal.</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623031, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.g004", "stats"=>{"downloads"=>0, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_Comparisons_of_the_infratemporal_ramus_of_the_jugal_in_basal_ceratopsians_/1623031", "title"=>"Comparisons of the infratemporal ramus of the jugal in basal ceratopsians.", "pos_in_sequence"=>0, "defined_type"=>1, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606244"], "description"=>"<p>(A-F) right quadrate and quadratojugal of <i>Hualianceratops wucaiwanensis</i>. (A) photograph in lateral view, (B) photograph in caudal view, (C) photograph in rostral view, (D) drawing in lateral view, (E) drawing in caudal view, (F) drawing in rostral view. (G-I) left quadrate of <i>Yinlong</i> (IVPP V18637) flipped for comparison with the right side. (G) Photograph in lateral view, (H) photograph in caudal view, (I) photograph in rostral view. Abbreviations: gr, groove; q, quadrate; qj, quadratojugal; qj.a, articulation with the quadratojugal; qn, quadrate notch.</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623032, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.g005", "stats"=>{"downloads"=>0, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_Comparison_of_the_quadrates_in_Hualianceratops_wucaiwanensis_and_Yinlong_downsi_/1623032", "title"=>"Comparison of the quadrates in <i>Hualianceratops wucaiwanensis</i> and <i>Yinlong downsi</i>.", "pos_in_sequence"=>0, "defined_type"=>1, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606245"], "description"=>"<p>(A) dorsal view, (B) ventral view, (C) left lateral view, (D) caudal view. Abbreviations: fo, fossa on the lateral surface; nd, nodes on the caudal margin; qp, stalked quadrate process.</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623033, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.g006", "stats"=>{"downloads"=>0, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_The_squamosal_of_Yinlong_downsi_IVPP_V18641_/1623033", "title"=>"The squamosal of <i>Yinlong downsi</i> (IVPP V18641).", "pos_in_sequence"=>0, "defined_type"=>1, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606246"], "description"=>"<p>(A) photograph in lateral view, (B) drawing in lateral view, (C) photograph in ventral view, (D) drawing in ventral view. Abbreviations: an, angular; d, dentary; emf, external mandibular fenestra; imf, inner mandibular fenestra; lsp, left splenial; pra, prearticular; pd, predentary; rsp, right splenial; sa, surangular.</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623034, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.g007", "stats"=>{"downloads"=>0, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_The_mandible_of_Hualianceratops_wucaiwanensis_IVPP_V18641_/1623034", "title"=>"The mandible of <i>Hualianceratops wucaiwanensis</i> (IVPP V18641).", "pos_in_sequence"=>0, "defined_type"=>1, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606247"], "description"=>"<p>(A) <i>Hualianceratops wucaiwanensis</i> (IVPP V18641), (B) <i>Hongshanosaurus houi</i> (IVPP V12617), (C) <i>Yinlong downsi</i> (IVPP V14530), (D) <i>Archaeoceratops oshimai</i> (IVPP V11114), (E) <i>Chaoyangsaurus youngi</i> (IGCAGS V371), (F) <i>Liaoceratops yanzigouensis</i> (IVPP V12633). Abbreviations: an, angular; co, coronoid; d, dentary; pd, predentary; sa, surangular.</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623035, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.g008", "stats"=>{"downloads"=>1, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_Comparison_of_the_mandible_in_basal_ceratopsians_in_right_lateral_view_/1623035", "title"=>"Comparison of the mandible in basal ceratopsians in right lateral view.", "pos_in_sequence"=>0, "defined_type"=>1, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606248"], "description"=>"<p>(A-B) two fused sacral vertebrates, cranial end is to the left. (A) ventral view, (B) dorsal view. (C) partial hind limb and left pes of <i>Hualianceratops wucaiwanensis</i>. Abbreviations: rf, right fibula; rt, right tibia; lf, left fibula; mt.I-II, metatarsal I-II; pII-IV, pes digits II-IV.</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623036, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.g009", "stats"=>{"downloads"=>0, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_Postcranial_material_of_Hualianceratops_wucaiwanensis_IVPP_V18641_/1623036", "title"=>"Postcranial material of <i>Hualianceratops wucaiwanensis</i> (IVPP V18641).", "pos_in_sequence"=>0, "defined_type"=>1, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606249"], "description"=>"<p>Bremer support (values above 1) is shown below and to the left of nodes, and bootstrap values are labeled to the left and above the nodes (values above 50%).</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623037, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.g010", "stats"=>{"downloads"=>2, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_Strict_consensus_tree_of_216_most_parsimonious_trees_produced_by_analysing_a_data_matrix_of_37_taxa_and_210_characters_/1623037", "title"=>"Strict consensus tree of 216 most parsimonious trees produced by analysing a data matrix of 37 taxa and 210 characters.", "pos_in_sequence"=>0, "defined_type"=>1, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606250"], "description"=>"<p>Ghost lineages (gray lines) implied by the stratigraphic distributions of taxa and the most parsimonious trees. Thick black lines indicate single occurrences, their length reflects uncertainty in dating. One of the phylogenies implying the fewest lineages is shown. Geochronology from [<a href=\"http://www.plosone.org/article/info:doi/10.1371/journal.pone.0143369#pone.0143369.ref041\" target=\"_blank\">41</a>].</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623038, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.g011", "stats"=>{"downloads"=>0, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_Ghost_lineages_implied_by_this_phylogenetic_analysis_/1623038", "title"=>"Ghost lineages implied by this phylogenetic analysis.", "pos_in_sequence"=>0, "defined_type"=>1, "published_date"=>"2015-12-09 11:56:56"}
  • {"files"=>["https://ndownloader.figshare.com/files/2606251"], "description"=>"<p>*estimated.</p><p>Skull measurements of <i>Hualianceratops</i> (IVPP 18641) and <i>Yinlong downsi</i> [IVPP 14530 (holotype), IVPP 18637], in millimeters.</p>", "links"=>[], "tags"=>["Hualianceratops wucaiwanensis gen", "subtemporal ramus", "quadrate", "Ceratopsia Ceratopsia", "phylogeny", "quadratojugal facet", "Hualianceratops shares", "novel hypothesis", "dentary", "ceratopsian lineages", "basal split", "ventral region", "Shishugou Formation", "basal ceratopsian", "sister group", "Xuanhuaceratop", "Chaoyangsauru", "Upper Jurassic", "Neoceratopsia", "rugose texturing", "Psittacosauru", "caudodorsal process", "basal ceratopsians Yinlong", "New Taxon", "Jurassic age", "herbivorous ornithischian clades", "Junggar Basin"], "article_id"=>1623039, "categories"=>["Biological Sciences"], "users"=>["Fenglu Han", "Catherine A. Forster", "James M. Clark", "Xing Xu"], "doi"=>"https://dx.doi.org/10.1371/journal.pone.0143369.t001", "stats"=>{"downloads"=>0, "page_views"=>0, "likes"=>0}, "figshare_url"=>"https://figshare.com/articles/_Skull_measurements_of_Hualianceratops_IVPP_18641_and_Yinlong_downsi_IVPP_14530_holotype_IVPP_18637_in_millimeters_/1623039", "title"=>"Skull measurements of <i>Hualianceratops</i> (IVPP 18641) and <i>Yinlong downsi</i> [IVPP 14530 (holotype), IVPP 18637], in millimeters.", "pos_in_sequence"=>0, "defined_type"=>3, "published_date"=>"2015-12-09 11:56:56"}

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Relative Metric

{"start_date"=>"2015-01-01T00:00:00Z", "end_date"=>"2015-12-31T00:00:00Z", "subject_areas"=>[]}
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