Selection for Social Signalling Drives the Evolution of Chameleon Colour Change
- Publication Date
- January 29, 2008
- Journal
- PLOS Biology
- Authors
- Devi Stuart Fox & Adnan Moussalli
- Volume
- 6
- Issue
- 1
- Pages
- e25
- DOI
- https://dx.plos.org/10.1371/journal.pbio.0060025
- Publisher URL
- http://journals.plos.org/plosbiology/article?id=10.1371%2Fjournal.pbio.0060025
- PubMed
- http://www.ncbi.nlm.nih.gov/pubmed/18232740
- PubMed Central
- http://www.ncbi.nlm.nih.gov/pmc/articles/PMC2214820
- Europe PMC
- http://europepmc.org/abstract/MED/18232740
- Web of Science
- 000254928300004
- Scopus
- 38949095386
- Mendeley
- http://www.mendeley.com/research/selection-social-signalling-drives-evolution-chameleon-colour-change-1
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Mendeley | Further Information
{"title"=>"Selection for social signalling drives the evolution of chameleon colour change", "type"=>"journal", "authors"=>[{"first_name"=>"Devi", "last_name"=>"Stuart-Fox", "scopus_author_id"=>"6602355314"}, {"first_name"=>"Adnan", "last_name"=>"Moussalli", "scopus_author_id"=>"6602290833"}], "year"=>2008, "source"=>"PLoS Biology", "identifiers"=>{"issn"=>"15449173", "scopus"=>"2-s2.0-38949095386", "sgr"=>"38949095386", "pui"=>"351226043", "isbn"=>"1544-9173", "pmid"=>"18232740", "doi"=>"10.1371/journal.pbio.0060025"}, "id"=>"4f867d23-9f96-3892-8033-178ce69b3379", "abstract"=>"Rapid colour change is a remarkable natural phenomenon that has evolved in several vertebrate and invertebrate lineages. The two principal explanations for the evolution of this adaptive strategy are (1) natural selection for crypsis (camouflage) against a range of different backgrounds and (2) selection for conspicuous social signals that maximise detectability to conspecifics, yet minimise exposure to predators because they are only briefly displayed. Here we show that evolutionary shifts in capacity for colour change in southern African dwarf chameleons (Bradypodion spp.) are associated with increasingly conspicuous signals used in male contests and courtship. To the chameleon visual system, species showing the most dramatic colour change display social signals that contrast most against the environmental background and amongst adjacent body regions. We found no evidence for the crypsis hypothesis, a finding reinforced by visual models of how both chameleons and their avian predators perceive chameleon colour variation. Instead, our results suggest that selection for conspicuous social signals drives the evolution of colour change in this system, supporting the view that transitory display traits should be under strong selection for signal detectability.", "link"=>"http://www.mendeley.com/research/selection-social-signalling-drives-evolution-chameleon-colour-change-1", "reader_count"=>266, "reader_count_by_academic_status"=>{"Professor > Associate Professor"=>14, "Student > Doctoral Student"=>13, "Researcher"=>55, "Student > Ph. D. Student"=>57, "Student > Postgraduate"=>13, "Student > Master"=>34, "Other"=>15, "Student > Bachelor"=>46, "Lecturer"=>3, "Lecturer > Senior Lecturer"=>2, "Professor"=>14}, "reader_count_by_user_role"=>{"Professor > Associate Professor"=>14, "Student > Doctoral Student"=>13, "Researcher"=>55, "Student > Ph. D. Student"=>57, "Student > Postgraduate"=>13, "Student > Master"=>34, "Other"=>15, "Student > Bachelor"=>46, "Lecturer"=>3, "Lecturer > Senior Lecturer"=>2, "Professor"=>14}, "reader_count_by_subject_area"=>{"Unspecified"=>9, "Agricultural and Biological Sciences"=>201, "Philosophy"=>1, "Chemistry"=>5, "Computer Science"=>2, "Earth and Planetary Sciences"=>7, "Economics, Econometrics and Finance"=>1, "Engineering"=>5, "Environmental Science"=>18, "Biochemistry, Genetics and Molecular Biology"=>5, "Nursing and Health Professions"=>1, "Materials Science"=>1, "Medicine and Dentistry"=>1, "Physics and Astronomy"=>4, "Psychology"=>4, "Social Sciences"=>1}, "reader_count_by_subdiscipline"=>{"Materials Science"=>{"Materials Science"=>1}, "Medicine and Dentistry"=>{"Medicine and Dentistry"=>1}, "Social Sciences"=>{"Social Sciences"=>1}, "Physics and Astronomy"=>{"Physics and Astronomy"=>4}, "Psychology"=>{"Psychology"=>4}, "Unspecified"=>{"Unspecified"=>9}, "Environmental Science"=>{"Environmental Science"=>18}, "Engineering"=>{"Engineering"=>5}, "Chemistry"=>{"Chemistry"=>5}, "Earth and Planetary Sciences"=>{"Earth and Planetary Sciences"=>7}, "Economics, Econometrics and Finance"=>{"Economics, Econometrics and Finance"=>1}, "Agricultural and Biological Sciences"=>{"Agricultural and Biological Sciences"=>201}, "Computer Science"=>{"Computer Science"=>2}, "Nursing and Health Professions"=>{"Nursing and Health Professions"=>1}, "Biochemistry, Genetics and Molecular Biology"=>{"Biochemistry, Genetics and Molecular Biology"=>5}, "Philosophy"=>{"Philosophy"=>1}}, "reader_count_by_country"=>{"Argentina"=>2, "Romania"=>1, "Hungary"=>1, "United States"=>12, "United Kingdom"=>2, "Portugal"=>2, "Switzerland"=>2, "Spain"=>3, "Costa Rica"=>1, "Canada"=>3, "Austria"=>1, "Brazil"=>4, "South Africa"=>4, "Italy"=>1, "Chile"=>1, "Germany"=>4}, "group_count"=>10}CrossRef
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Wikipedia | Further Information
- {"title"=>"Chameleon", "url"=>"http://en.wikipedia.org/wiki/Chameleon", "timestamp"=>"2018-11-18T06:16:36Z"}
- {"title"=>"Ecology", "url"=>"http://en.wikipedia.org/wiki/Ecology", "timestamp"=>"2018-11-13T20:02:40Z"}
- {"title"=>"Camaleão", "url"=>"http://pt.wikipedia.org/wiki/Camaleão", "timestamp"=>"2018-09-08T13:35:04Z"}
- {"title"=>"Cromatóforo", "url"=>"http://pt.wikipedia.org/wiki/Cromatóforo", "timestamp"=>"2018-07-29T11:38:09Z"}
- {"title"=>"Kameleoner", "url"=>"http://no.wikipedia.org/wiki/Kameleoner", "timestamp"=>"2018-05-16T19:41:30Z"}
- {"title"=>"File:Bradypodion atromontanum1.jpg", "url"=>"http://commons.wikimedia.org/wiki/File:Bradypodion_atromontanum1.jpg", "timestamp"=>"2012-10-01T09:47:50Z"}
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- {"month"=>"1", "year"=>"2010", "pdf_views"=>"18", "xml_views"=>"4", "html_views"=>"193"}
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- {"month"=>"4", "year"=>"2010", "pdf_views"=>"38", "xml_views"=>"1", "html_views"=>"227"}
- {"month"=>"5", "year"=>"2010", "pdf_views"=>"29", "xml_views"=>"0", "html_views"=>"219"}
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- {"month"=>"8", "year"=>"2010", "pdf_views"=>"16", "xml_views"=>"0", "html_views"=>"124"}
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- {"month"=>"10", "year"=>"2010", "pdf_views"=>"54", "xml_views"=>"1", "html_views"=>"265"}
- {"month"=>"11", "year"=>"2010", "pdf_views"=>"60", "xml_views"=>"0", "html_views"=>"390"}
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- {"month"=>"4", "year"=>"2011", "pdf_views"=>"35", "xml_views"=>"2", "html_views"=>"304"}
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- {"month"=>"4", "year"=>"2012", "pdf_views"=>"37", "xml_views"=>"1", "html_views"=>"281"}
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- {"month"=>"6", "year"=>"2012", "pdf_views"=>"28", "xml_views"=>"2", "html_views"=>"129"}
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- {"month"=>"8", "year"=>"2012", "pdf_views"=>"10", "xml_views"=>"6", "html_views"=>"134"}
- {"month"=>"9", "year"=>"2012", "pdf_views"=>"26", "xml_views"=>"0", "html_views"=>"159"}
- {"month"=>"10", "year"=>"2012", "pdf_views"=>"49", "xml_views"=>"0", "html_views"=>"280"}
- {"month"=>"11", "year"=>"2012", "pdf_views"=>"41", "xml_views"=>"0", "html_views"=>"302"}
- {"month"=>"12", "year"=>"2012", "pdf_views"=>"20", "xml_views"=>"0", "html_views"=>"212"}
- {"month"=>"1", "year"=>"2013", "pdf_views"=>"21", "xml_views"=>"0", "html_views"=>"250"}
- {"month"=>"2", "year"=>"2013", "pdf_views"=>"31", "xml_views"=>"3", "html_views"=>"453"}
- {"month"=>"3", "year"=>"2013", "pdf_views"=>"27", "xml_views"=>"0", "html_views"=>"416"}
- {"month"=>"4", "year"=>"2013", "pdf_views"=>"45", "xml_views"=>"0", "html_views"=>"475"}
- {"month"=>"5", "year"=>"2013", "pdf_views"=>"27", "xml_views"=>"0", "html_views"=>"418"}
- {"month"=>"6", "year"=>"2013", "pdf_views"=>"24", "xml_views"=>"4", "html_views"=>"282"}
- {"month"=>"7", "year"=>"2013", "pdf_views"=>"21", "xml_views"=>"1", "html_views"=>"151"}
- {"month"=>"8", "year"=>"2013", "pdf_views"=>"14", "xml_views"=>"0", "html_views"=>"179"}
- {"month"=>"9", "year"=>"2013", "pdf_views"=>"25", "xml_views"=>"0", "html_views"=>"388"}
- {"month"=>"10", "year"=>"2013", "pdf_views"=>"42", "xml_views"=>"1", "html_views"=>"369"}
- {"month"=>"11", "year"=>"2013", "pdf_views"=>"54", "xml_views"=>"4", "html_views"=>"448"}
- {"month"=>"12", "year"=>"2013", "pdf_views"=>"21", "xml_views"=>"2", "html_views"=>"308"}
- {"month"=>"1", "year"=>"2014", "pdf_views"=>"32", "xml_views"=>"0", "html_views"=>"316"}
- {"month"=>"2", "year"=>"2014", "pdf_views"=>"32", "xml_views"=>"0", "html_views"=>"313"}
- {"month"=>"3", "year"=>"2014", "pdf_views"=>"41", "xml_views"=>"2", "html_views"=>"378"}
- {"month"=>"4", "year"=>"2014", "pdf_views"=>"48", "xml_views"=>"1", "html_views"=>"465"}
- {"month"=>"5", "year"=>"2014", "pdf_views"=>"31", "xml_views"=>"2", "html_views"=>"299"}
- {"month"=>"6", "year"=>"2014", "pdf_views"=>"16", "xml_views"=>"0", "html_views"=>"205"}
- {"month"=>"7", "year"=>"2014", "pdf_views"=>"20", "xml_views"=>"0", "html_views"=>"192"}
- {"month"=>"8", "year"=>"2014", "pdf_views"=>"18", "xml_views"=>"2", "html_views"=>"140"}
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- {"month"=>"11", "year"=>"2014", "pdf_views"=>"28", "xml_views"=>"2", "html_views"=>"455"}
- {"month"=>"12", "year"=>"2014", "pdf_views"=>"11", "xml_views"=>"3", "html_views"=>"254"}
- {"month"=>"1", "year"=>"2015", "pdf_views"=>"18", "xml_views"=>"0", "html_views"=>"212"}
- {"month"=>"2", "year"=>"2015", "pdf_views"=>"23", "xml_views"=>"1", "html_views"=>"262"}
- {"month"=>"3", "year"=>"2015", "pdf_views"=>"36", "xml_views"=>"0", "html_views"=>"363"}
- {"month"=>"4", "year"=>"2015", "pdf_views"=>"30", "xml_views"=>"1", "html_views"=>"341"}
- {"month"=>"5", "year"=>"2015", "pdf_views"=>"20", "xml_views"=>"0", "html_views"=>"287"}
- {"month"=>"6", "year"=>"2015", "pdf_views"=>"12", "xml_views"=>"0", "html_views"=>"199"}
- {"month"=>"7", "year"=>"2015", "pdf_views"=>"14", "xml_views"=>"0", "html_views"=>"120"}
- {"month"=>"8", "year"=>"2015", "pdf_views"=>"8", "xml_views"=>"0", "html_views"=>"127"}
- {"month"=>"9", "year"=>"2015", "pdf_views"=>"35", "xml_views"=>"0", "html_views"=>"198"}
- {"month"=>"10", "year"=>"2015", "pdf_views"=>"24", "xml_views"=>"0", "html_views"=>"297"}
- {"month"=>"11", "year"=>"2015", "pdf_views"=>"42", "xml_views"=>"0", "html_views"=>"325"}
- {"month"=>"12", "year"=>"2015", "pdf_views"=>"18", "xml_views"=>"0", "html_views"=>"220"}
- {"month"=>"1", "year"=>"2016", "pdf_views"=>"34", "xml_views"=>"0", "html_views"=>"185"}
- {"month"=>"2", "year"=>"2016", "pdf_views"=>"55", "xml_views"=>"0", "html_views"=>"231"}
- {"month"=>"3", "year"=>"2016", "pdf_views"=>"45", "xml_views"=>"0", "html_views"=>"283"}
- {"month"=>"4", "year"=>"2016", "pdf_views"=>"27", "xml_views"=>"0", "html_views"=>"262"}
- {"month"=>"5", "year"=>"2016", "pdf_views"=>"13", "xml_views"=>"0", "html_views"=>"156"}
- {"month"=>"6", "year"=>"2016", "pdf_views"=>"16", "xml_views"=>"0", "html_views"=>"136"}
- {"month"=>"7", "year"=>"2016", "pdf_views"=>"11", "xml_views"=>"0", "html_views"=>"112"}
- {"month"=>"8", "year"=>"2016", "pdf_views"=>"25", "xml_views"=>"0", "html_views"=>"138"}
- {"month"=>"9", "year"=>"2016", "pdf_views"=>"35", "xml_views"=>"0", "html_views"=>"180"}
- {"month"=>"10", "year"=>"2016", "pdf_views"=>"36", "xml_views"=>"0", "html_views"=>"207"}
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- {"month"=>"12", "year"=>"2016", "pdf_views"=>"69", "xml_views"=>"1", "html_views"=>"329"}
- {"month"=>"1", "year"=>"2017", "pdf_views"=>"23", "xml_views"=>"2", "html_views"=>"319"}
- {"month"=>"2", "year"=>"2017", "pdf_views"=>"53", "xml_views"=>"1", "html_views"=>"530"}
- {"month"=>"3", "year"=>"2017", "pdf_views"=>"47", "xml_views"=>"2", "html_views"=>"490"}
- {"month"=>"4", "year"=>"2017", "pdf_views"=>"38", "xml_views"=>"1", "html_views"=>"750"}
- {"month"=>"5", "year"=>"2017", "pdf_views"=>"94", "xml_views"=>"1", "html_views"=>"484"}
- {"month"=>"6", "year"=>"2017", "pdf_views"=>"81", "xml_views"=>"0", "html_views"=>"184"}
- {"month"=>"7", "year"=>"2017", "pdf_views"=>"33", "xml_views"=>"0", "html_views"=>"178"}
- {"month"=>"8", "year"=>"2017", "pdf_views"=>"35", "xml_views"=>"1", "html_views"=>"138"}
- {"month"=>"9", "year"=>"2017", "pdf_views"=>"116", "xml_views"=>"2", "html_views"=>"170"}
- {"month"=>"10", "year"=>"2017", "pdf_views"=>"199", "xml_views"=>"3", "html_views"=>"368"}
- {"month"=>"11", "year"=>"2017", "pdf_views"=>"112", "xml_views"=>"1", "html_views"=>"358"}
- {"month"=>"12", "year"=>"2017", "pdf_views"=>"16", "xml_views"=>"1", "html_views"=>"220"}
- {"month"=>"1", "year"=>"2018", "pdf_views"=>"45", "xml_views"=>"0", "html_views"=>"252"}
- {"month"=>"2", "year"=>"2018", "pdf_views"=>"44", "xml_views"=>"0", "html_views"=>"162"}
- {"month"=>"3", "year"=>"2018", "pdf_views"=>"49", "xml_views"=>"2", "html_views"=>"193"}
- {"month"=>"4", "year"=>"2018", "pdf_views"=>"46", "xml_views"=>"1", "html_views"=>"217"}
- {"month"=>"5", "year"=>"2018", "pdf_views"=>"35", "xml_views"=>"0", "html_views"=>"131"}
- {"month"=>"6", "year"=>"2018", "pdf_views"=>"15", "xml_views"=>"0", "html_views"=>"54"}
- {"month"=>"7", "year"=>"2018", "pdf_views"=>"19", "xml_views"=>"5", "html_views"=>"55"}
- {"month"=>"8", "year"=>"2018", "pdf_views"=>"29", "xml_views"=>"2", "html_views"=>"57"}
- {"month"=>"9", "year"=>"2018", "pdf_views"=>"39", "xml_views"=>"2", "html_views"=>"104"}
- {"month"=>"10", "year"=>"2018", "pdf_views"=>"89", "xml_views"=>"3", "html_views"=>"220"}
- {"month"=>"11", "year"=>"2018", "pdf_views"=>"44", "xml_views"=>"1", "html_views"=>"169"}
- {"month"=>"12", "year"=>"2018", "pdf_views"=>"38", "xml_views"=>"2", "html_views"=>"94"}
- {"month"=>"1", "year"=>"2019", "pdf_views"=>"22", "xml_views"=>"0", "html_views"=>"157"}
- {"month"=>"2", "year"=>"2019", "pdf_views"=>"47", "xml_views"=>"0", "html_views"=>"182"}
- {"month"=>"3", "year"=>"2019", "pdf_views"=>"74", "xml_views"=>"1", "html_views"=>"225"}
- {"month"=>"4", "year"=>"2019", "pdf_views"=>"50", "xml_views"=>"2", "html_views"=>"139"}
- {"month"=>"5", "year"=>"2019", "pdf_views"=>"46", "xml_views"=>"1", "html_views"=>"95"}
- {"month"=>"6", "year"=>"2019", "pdf_views"=>"17", "xml_views"=>"0", "html_views"=>"56"}
- {"month"=>"7", "year"=>"2019", "pdf_views"=>"31", "xml_views"=>"1", "html_views"=>"22"}
- {"month"=>"8", "year"=>"2019", "pdf_views"=>"16", "xml_views"=>"0", "html_views"=>"45"}
- {"month"=>"9", "year"=>"2019", "pdf_views"=>"49", "xml_views"=>"0", "html_views"=>"113"}
- {"month"=>"10", "year"=>"2019", "pdf_views"=>"33", "xml_views"=>"1", "html_views"=>"143"}
- {"month"=>"11", "year"=>"2019", "pdf_views"=>"53", "xml_views"=>"1", "html_views"=>"155"}
- {"month"=>"12", "year"=>"2019", "pdf_views"=>"88", "xml_views"=>"0", "html_views"=>"133"}
- {"month"=>"1", "year"=>"2020", "pdf_views"=>"105", "xml_views"=>"1", "html_views"=>"89"}
- {"month"=>"2", "year"=>"2020", "pdf_views"=>"217", "xml_views"=>"0", "html_views"=>"58"}
- {"month"=>"3", "year"=>"2020", "pdf_views"=>"156", "xml_views"=>"3", "html_views"=>"140"}
- {"month"=>"4", "year"=>"2020", "pdf_views"=>"150", "xml_views"=>"1", "html_views"=>"59"}
- {"month"=>"5", "year"=>"2020", "pdf_views"=>"169", "xml_views"=>"0", "html_views"=>"78"}
- {"month"=>"6", "year"=>"2020", "pdf_views"=>"71", "xml_views"=>"2", "html_views"=>"45"}
- {"month"=>"7", "year"=>"2020", "pdf_views"=>"26", "xml_views"=>"0", "html_views"=>"27"}
- {"month"=>"8", "year"=>"2020", "pdf_views"=>"34", "xml_views"=>"0", "html_views"=>"59"}
- {"month"=>"9", "year"=>"2020", "pdf_views"=>"67", "xml_views"=>"3", "html_views"=>"140"}
- {"month"=>"10", "year"=>"2020", "pdf_views"=>"81", "xml_views"=>"3", "html_views"=>"163"}
- {"month"=>"11", "year"=>"2020", "pdf_views"=>"170", "xml_views"=>"3", "html_views"=>"200"}
- {"month"=>"12", "year"=>"2020", "pdf_views"=>"60", "xml_views"=>"1", "html_views"=>"102"}
- {"month"=>"1", "year"=>"2021", "pdf_views"=>"22", "xml_views"=>"0", "html_views"=>"60"}
- {"month"=>"2", "year"=>"2021", "pdf_views"=>"29", "xml_views"=>"2", "html_views"=>"133"}
PMC Usage Stats | Further Information
- {"scanned-page-browse"=>"0", "month"=>"1", "cited-by"=>"0", "abstract"=>"0", "full-text"=>"47", "unique-ip"=>"38", "pdf"=>"0", "year"=>"2010", "figure"=>"1", "scanned-summary"=>"0", "supp-data"=>"0"}
- {"month"=>"2", "scanned-page-browse"=>"0", "cited-by"=>"0", "abstract"=>"0", "full-text"=>"48", "year"=>"2010", "pdf"=>"0", "unique-ip"=>"37", "figure"=>"1", "scanned-summary"=>"0", "supp-data"=>"0"}
- {"scanned-page-browse"=>"0", "month"=>"3", "cited-by"=>"1", "abstract"=>"0", "full-text"=>"196", "unique-ip"=>"60", "pdf"=>"0", "year"=>"2010", "figure"=>"28", "scanned-summary"=>"0", "supp-data"=>"1"}
- {"month"=>"4", "scanned-page-browse"=>"0", "cited-by"=>"0", "abstract"=>"1", "full-text"=>"126", "year"=>"2010", "pdf"=>"0", "unique-ip"=>"68", "figure"=>"7", "scanned-summary"=>"0", "supp-data"=>"2"}
- {"scanned-page-browse"=>"0", "month"=>"5", "cited-by"=>"0", "abstract"=>"2", "full-text"=>"120", "unique-ip"=>"69", "pdf"=>"0", "year"=>"2010", "figure"=>"5", "scanned-summary"=>"0", "supp-data"=>"0"}
- {"month"=>"6", "scanned-page-browse"=>"0", "cited-by"=>"1", "abstract"=>"4", "full-text"=>"77", "year"=>"2010", "pdf"=>"0", "unique-ip"=>"44", "figure"=>"11", "scanned-summary"=>"0", "supp-data"=>"1"}
- {"scanned-page-browse"=>"0", "month"=>"7", "cited-by"=>"0", "abstract"=>"0", "full-text"=>"25", "unique-ip"=>"15", "pdf"=>"0", "year"=>"2010", "figure"=>"1", "scanned-summary"=>"0", "supp-data"=>"0"}
- {"month"=>"8", "scanned-page-browse"=>"0", "cited-by"=>"0", "abstract"=>"0", "full-text"=>"34", "year"=>"2010", "pdf"=>"0", "unique-ip"=>"22", "figure"=>"1", "scanned-summary"=>"0", "supp-data"=>"0"}
- {"scanned-page-browse"=>"0", "month"=>"9", "cited-by"=>"0", "abstract"=>"0", "full-text"=>"64", "unique-ip"=>"47", "pdf"=>"0", "year"=>"2010", "figure"=>"1", "scanned-summary"=>"0", "supp-data"=>"2"}
- {"month"=>"10", "scanned-page-browse"=>"0", "cited-by"=>"0", "abstract"=>"2", "full-text"=>"50", "year"=>"2010", "pdf"=>"0", "unique-ip"=>"41", "figure"=>"16", "scanned-summary"=>"0", "supp-data"=>"2"}
- {"scanned-page-browse"=>"0", "month"=>"11", "cited-by"=>"0", "abstract"=>"0", "full-text"=>"76", "unique-ip"=>"44", "pdf"=>"0", "year"=>"2010", "figure"=>"21", "scanned-summary"=>"0", "supp-data"=>"5"}
- {"month"=>"12", "scanned-page-browse"=>"0", "cited-by"=>"0", "abstract"=>"0", "full-text"=>"47", "year"=>"2010", "pdf"=>"0", "unique-ip"=>"38", "figure"=>"12", "scanned-summary"=>"0", "supp-data"=>"0"}
- {"scanned-page-browse"=>"0", "month"=>"1", "cited-by"=>"0", "abstract"=>"2", "full-text"=>"42", "unique-ip"=>"32", "pdf"=>"0", "year"=>"2011", "figure"=>"2", "scanned-summary"=>"0", "supp-data"=>"0"}
- {"month"=>"2", "scanned-page-browse"=>"0", "cited-by"=>"0", "abstract"=>"0", "full-text"=>"42", "year"=>"2011", "pdf"=>"0", "unique-ip"=>"37", "figure"=>"3", "scanned-summary"=>"0", "supp-data"=>"3"}
- {"scanned-page-browse"=>"0", "month"=>"3", "cited-by"=>"2", "abstract"=>"2", "full-text"=>"74", "unique-ip"=>"37", "pdf"=>"0", "year"=>"2011", "figure"=>"6", "scanned-summary"=>"0", "supp-data"=>"2"}
- {"month"=>"4", "scanned-page-browse"=>"0", "cited-by"=>"0", "abstract"=>"1", "full-text"=>"68", "year"=>"2011", "pdf"=>"0", "unique-ip"=>"51", "figure"=>"3", "scanned-summary"=>"0", "supp-data"=>"2"}
- {"scanned-page-browse"=>"0", "month"=>"5", "cited-by"=>"2", "abstract"=>"3", "full-text"=>"52", "unique-ip"=>"39", "pdf"=>"0", "year"=>"2011", "figure"=>"5", "scanned-summary"=>"0", "supp-data"=>"0"}
- {"month"=>"6", "scanned-page-browse"=>"0", "cited-by"=>"0", "abstract"=>"1", "full-text"=>"45", "year"=>"2011", "pdf"=>"0", "unique-ip"=>"30", "figure"=>"15", "scanned-summary"=>"0", "supp-data"=>"0"}
- {"scanned-page-browse"=>"0", "month"=>"7", "cited-by"=>"1", "abstract"=>"0", "full-text"=>"25", "unique-ip"=>"25", "pdf"=>"0", "year"=>"2011", "figure"=>"1", "scanned-summary"=>"0", "supp-data"=>"0"}
- {"month"=>"8", "scanned-page-browse"=>"0", "cited-by"=>"0", "abstract"=>"0", "full-text"=>"29", "year"=>"2011", "pdf"=>"0", "unique-ip"=>"23", "figure"=>"4", "scanned-summary"=>"0", "supp-data"=>"0"}
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Relative Metric
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